Synchronous update revisited
Directed Paths in Random Media (DPRM): Least energy paths obtained recursively (transfer matrix, dynamic programming)
Synchronous growth with variable size is equivalent to DPRM leads to similar statistics of sectors and ancestry:
"Evolution in range expansions with competition at rough boundaries,"
S. Chu, M.Kardar, D.R. Nelson, & D.A. Beller, Journal of Theoretical Biology 478, 153 (2019). (off-line)
Fitness and Mutation may serve as an alternative interpretation for optimal energy and random landscape:
What is the distribution of fitness at the front of range expansion?
The stepping stone model predics a (non-Gaussian) Tracy-Widom distribution:
Different fitness distributions are likely for different areas of competition:
"The Mother Machine"
versus